X chromosome drive in a widespread Palearctic woodland fly, Drosophila testacea

dc.contributor.authorKeais, Graeme
dc.contributor.authorHanson, Mark
dc.contributor.authorGowen, Brent
dc.contributor.authorPerlman, Steve
dc.date.accessioned2017-07-31T17:24:28Z
dc.date.available2017-07-31T17:24:28Z
dc.date.copyright2017en_US
dc.date.issued2017-06
dc.description.abstractSelfish genes that bias their own transmission during meiosis can spread rapidly in populations, even if they contribute negatively to the fitness of their host. Driving X chromosomes provide a clear example of this type of selfish propagation. These chromosomes have important evolutionary and ecological consequences, and can be found in a broad range of taxa including plants, mammals and insects. Here, we report a new case of X chromosome drive (X drive) in a widespread woodland fly, Drosophila testacea. We show that males carrying the driving X (SR males) sire 80–100% female offspring and possess a diagnostic X chromosome haplotype that is perfectly associated with the sex ratio distortion phenotype. We find that the majority of sons produced by SR males are sterile and appear to lack a Y chromosome, suggesting that meiotic defects involving the Y chromosome may underlie X drive in this species. Abnormalities in sperm cysts of SR males reflect that some spermatids are failing to develop properly, confirming that drive is acting during gametogenesis. By screening wild-caught flies using progeny sex ratios and a diagnostic marker, we demonstrate that the driving X is present in wild populations at a frequency of ~ 10% and that suppressors of drive are segregating in the same population. The testacea species group appears to be a hot spot for X drive, and D. testacea is a promising model to compare driving X chromosomes in closely related species, some of which may even be younger than the chromosomes themselves.en_US
dc.description.reviewstatusRevieweden_US
dc.description.scholarlevelFacultyen_US
dc.identifier.citationKeais, G.L., Hanson, M.A., Gowen, B.E., Perlman, S.J. (2017). X chromosome drive in a widespread Palearctic woodland fly, Drosophila testacea. Journal of Evolutionary Biology, 30(6), 1185-1194. doi: 10.1111/jeb.13089en_US
dc.identifier.urihttps://doi.org/10.1111/jeb.13089
dc.identifier.urihttp://hdl.handle.net/1828/8385
dc.language.isoenen_US
dc.publisherJournal of Evolutionary Biologyen_US
dc.subjectDrosophila
dc.subjectgenetic conflict
dc.subjectmeiotic drive
dc.subjectsegregation distortion
dc.subjectselfish genetic elements
dc.subjectX chromosome drive
dc.subject.departmentDepartment of Biology
dc.titleX chromosome drive in a widespread Palearctic woodland fly, Drosophila testaceaen_US
dc.typePreprinten_US

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